Omphalos: An Attempt to Untie the Geological Knot. Gosse Philip Henry

the Reptiles of the Eocene beds, exhibit a great preponderance of agreement with those that now exist, so far as genus is concerned, though the species are still almost wholly distinct. The approximation is particularly marked in the Molluscous sub-kingdom, by the almost entire disappearance of the hitherto swarming Brachiopod and Cephalopod forms, and the progressive substitution for them of the Conchifera and Gastropoda, which had, however, throughout the Secondary epoch, been gradually coming forward to their present predominance in nature.

      Among the Fishes, the Placoid type was diminished in number; and those that were produced were mostly Sharks and Rays, of modern genera; but the chief difference was the paucity of those mailed forms (Ganoids), which were so abundant during the Oolitic period. On the other hand, the Ctenoid and Cycloid forms, which had begun to make their appearance in small numbers in the Chalk, are well represented. In both this deficiency and this plenitude, there is a very decided approach to existing conditions; for the Ganoids are almost unknown with us, while the last-named two orders are abundant. Representatives of our Perches, Maigres, Mackerels, Blennies, Herrings, and Cods, were numerous; distinct, however, from the present species. But not a single member of the great Salmon family was yet introduced.

      The great Saurian Reptiles had entirely disappeared, and were quite unrepresented in the tertiary beds, except by a Crocodile or two, and a small Lizard. Turtles were, however, numerous, both of the marine and lacustrine kinds; and there is an interesting stranger, in the form of a large Serpent, allied to our Pythons, some twenty feet in length.

      Birds and Mammals began now to assume their place on the land. The London Clay presents us with a little Vulture; and the Paris basin contains remnants of species representing the Raptores, the Rasores, the Grallatores, and the Natatores.

      The Quadrupeds came in in some force; not developed from the lowest to the highest scale of organization; for the Monkey and the Bat occur in sands, certainly not later, if not earlier, than the London Clay, contemporaneously with the Racoon, and before the existence of any Rodent or Cetacean. Some Carnivora, as the Wolf and the Fox, roamed the woods, but the character of the epoch was given by the Pachyderms.

      These, however, were not the massive colossi that browse in the African or Indian jungles of our days; no Elephant, no Rhinoceros, no Hippopotamus was as yet formed. But several kinds of Tapir wallowed in the morasses; and a goodly number of largish beasts, whose affinities were with the Pachydermata, while their analogies were with the Ruminantia, served as substitutes for the latter order, which was wholly wanting. These interesting quadrupeds, forming the genus Anoplotherium, were remarkable for two peculiarities, – their feet were two-toed, and their teeth were ranged in a continuous series, without any interval between the incisors and the molars. They varied in size from that of an ass to that of a hare.

      The physical conditions of our earth, when it was tenanted by these creatures, is thus described: – "All the great plains of Europe, and the districts through which the principal rivers now run, were then submerged; in all probability, the land chiefly extended in a westerly direction, far out into the Atlantic, possibly trending to the south, and connecting the western shores of England with the volcanic islands off the west coast of Africa. The great mountain chains of Europe, the Pyrenees, the Alps, the Apennines, the mountains of Greece, the mountains of Bohemia, and the Carpathians, existed then only as chains of islands in an open sea. Elevatory movements, having an east and west direction, had, however, already commenced, and were producing important results, laying bare the Wealden district in the south-east of England. The southern and central European district, and parts of western Asia, were the recipients of calcareous deposits (chiefly the skeletons of Foraminifera), forming the Apennine limestone; while numerous islands were gradually lifted above the sea, and fragments of disturbed and fractured rock were washed upon the neighbouring shallows or coast-lines, forming beds of gravel covering the Chalk. The beds of Nummulites and Miliolites, contemporaneous with those containing the Sheppey plants and the Paris quadrupeds, seem to indicate a deep sea at no great distance from shore, and render it probable that there were frequent alternations of elevation and depression, perhaps the result of disturbances acting in the direction already alluded to.

      "The shores of the islands and main land were, however, occasionally low and swampy, rivers bringing down mud in what is now the south-east of England, and the neighbourhood of Brussels, but depositing extensive calcareous beds near Paris. Deep inlets of the sea, estuaries, and the shifting mouths of a river, were also affected by numerous alterations of level not sufficient to destroy, but powerful enough to modify, the animal and vegetable species then existing; and these movements were continued for a long time."35

      After the elevation of the mountain summits of Europe above the sea, and while the same causes were still in operation, deposits were being made in the narrow intervening seas of the Archipelago, such as the present south of France, the valleys of the Rhine and Danube, the eastern districts of England and Portugal. These deposits were partly marine and partly lacustrine; the former consisting largely of loose sands, mingled with shells and gravel. In Switzerland is a thick mass of conglomerate; and in the district around Mayence, there is a series of fresh-water limestones, and sandstones charged with organic remains.

      The changes which took place during this comparatively recent epoch were not sudden, but gradual; the results of operations which were probably going on without intermission, and perhaps have not yet ceased. The land was more and more upheaved, till at length, what had been an archipelago of islands became a continent, and Europe assumed the form which it bears on our maps.

      The most interesting addition to the natural history of the Miocene, or Middle Tertiary period, was the Dinotherium– a huge Pachyderm, twice as large as an elephant, with a tapir-like proboscis, and two great tusks curving downward from the lower jaw. It was, doubtless, aquatic in its habits, and possibly (for its hinder parts are not known), it may have been allied to the Dugong and Manatee, those whale-like Pachyderms, with a broad horizontal tail, instead of posterior limbs.

      Other great herbivorous beasts roamed over the new-made land. The Mastodons, closely allied to the Elephant, had their head-quarters in North America, but extended also to Europe. And the Elephants themselves, of several species, were spread over the northern hemisphere, even to the polar regions. The Hippopotamus, the Rhinoceros, and other creatures, now exclusively tropical, were also inhabitants of the same northern latitudes.

      From some specimens of Elephants and Rhinoceroses of this period, which seem to have been buried in avalanches, and thus to have been preserved from decomposition, even of the more transitory parts, as muscle and skin, we learn something of the climate that prevailed. The very fact of their preservation, by the antiseptic power of frost, shows that it was not a tropical climate in which they lived; and the clothing of thick wool, fur, and hair, which protected the skin of the Mammoth, or Siberian Elephant, tends to the same conclusion. At the same time, those regions were not so intensely cold as they are now. For the district in which the remains of Elephants and their associates are found, in almost incredible abundance, is that inhospitable coast of northern Asia which bounds the Polar Sea.

      The trees of a temperate climate – the oak, the beech, the maple, the poplar, and the birch – which now attain their highest limit somewhere about 70° of north latitude, and there are dwarfed to minute shrubs, appear then to have grown at the very verge of the polar basin; and that in the condition of vast and luxuriant forests, perhaps occupying sheltered valleys between mountains whose steep sides were covered with snow, already become perennial, and ever and anon rolling down in overwhelming avalanches, such as those which now occasionally descend into the valleys of the Swiss Alps.

      The coast of Suffolk displays a formation known as the Crag – a local name for gravel – which rests partly on the chalk; but, as it lies in other parts over the London Clay, it is assigned to the later Tertiary, or what is called the Pleiocene period. It is divided into the coralline and the red crag, the latter being uppermost where they exist together, and therefore being the more recent. The Coralline Crag is nearly composed of corals and shells, the former almost wholly extinct now; but the latter containing upwards of seventy species still existing in the adjacent seas. The Red Crag contains few zoophytes, but is remarkable for the remains of at least five species of Whales. Other Mammalia occur in this formation,

35

Ansted's Anc. World, 267.