More Letters of Charles Darwin — Volume 1. Darwin Charles

by Mr. Darwin for future reference in the third and fourth editions, and have been only able to find the following, which occurs in the third edition (Volume I., 1836, page 242) (14/8. There is also (ed. 1837, Volume II., page 344) a vague reference to Natural Selection, of which the last sentence is enclosed in pencil in inverted commas, as though Mr. Darwin had intended to quote it: "In other parts of Africa the xanthous variety {of man} often appears, but does not multiply. Individuals thus characterised are like seeds which perish in an uncongenial soil.")

      "The variety in form, prevalent among all organised productions of nature, is found to subsist between individual beings of whatever species, even when they are offspring of the same parents. Another circumstance equally remarkable is the tendency which exists in almost every tribe, whether of animals or of plants, to transmit to their offspring and to perpetuate in their race all individual peculiarities which may thus have taken their rise. These two general facts in the economy of organised beings lay a foundation for the existence of diversified races, originating from the same primitive stock and within the limits of identical species."

      On the following page (page 243) a passage (not marked by Mr. Darwin) emphasises the limitation which Prichard ascribed to the results of variation and inheritance: —

      "Even those physiologists who contend for what is termed the indefinite nature of species admit that they have limits at present and under ordinary circumstances. Whatever diversities take place happen without breaking in upon the characteristic type of the species. This is transmitted from generation to generation: goats produce goats, and sheep, sheep."

      The passage on page 242 occurs in the reprint of the 1836-7 edition which forms part of the 1841-51 edition, but is not there marked by Mr. Darwin. He notes at the end of Volume I. of the 1836-7 edition: "March, 1857. I have not looked through all these {i.e. marked passages}, but I have gone through the later edition"; and a similar entry is in Volume II. of the third edition. It is therefore easy to understand how he came to overlook the passage on page 242 when he began the fuller statement of his species theory which is referred to in the "Life and Letters" as the "unfinished book." In the historical sketch prefixed to the "Origin of Species" writers are named as precursors whose claims are less strong than Prichard's, and it is certain that Mr. Darwin would have given an account of him if he had thought of him as an evolutionist.

      The two following passages will show that Mr. Darwin was, from his knowledge of Prichard's books, justified in classing him among those who did not believe in the mutability of species:

      "The various tribes of organised beings were originally placed by the Creator in certain regions, for which they are by their nature peculiarly adapted. Each species had only one beginning in a single stock: probably a single pair, as Linnaeus supposed, was first called into being in some particular spot, and the progeny left to disperse themselves to as great a distance from the original centre of their existence as the locomotive powers bestowed on them, or their capability of bearing changes of climate and other physical agencies, may have enabled them to wander." (14/9. Prichard, third edition, 1836-7, Volume I., page 96.)

      The second passage is annotated by Mr. Darwin with a shower of exclamation marks:

      "The meaning attached to the term SPECIES in natural history is very definite and intelligible. It includes only the following conditions — namely, separate origin and distinctness of race, evinced by the constant transmission of some characteristic peculiarity of organisation. A race of animals or of plants marked by any peculiar character which has always been constant and undeviating constitutes a species; and two races are considered as specifically different, if they are distinguished from each other by some characteristic which one cannot be supposed to have acquired, or the other to have lost through any known operation of physical causes; for we are hence led to conclude that the tribes thus distinguished have not descended from the same original stock." (14/10. Prichard, ed. 1836-7, Volume I., page 106. This passage is almost identical with that quoted from the second edition, Volume I., page 90. The latter part, from "and two races..." occurs in the second edition, though not quoted above.)

      As was his custom, Mr. Darwin pinned at the end of the first volume of the 1841-51 edition a piece of paper containing a list of the pages where marked passages occur. This paper bears, written in pencil, "How like my book all this will be!" The words appear to refer to Prichard's discussion on the dispersal of animals and plants; they certainly do not refer to the evolutionary views to be found in the book.)

      LETTER 15. TO J.D. HOOKER. Down {1844}.

      Thank you exceedingly for your long letter, and I am in truth ashamed of the time and trouble you have taken for me; but I must some day write again to you on the subject of your letter. I will only now observe that you have extended my remark on the range of species of shells into the range of genera or groups. Analogy from shells would only go so far, that if two or three species...were found to range from America to India, they would be found to extend through an unusual thickness of strata — say from the Upper Cretaceous to its lowest bed, or the Neocomian. Or you may reverse it and say those species which range throughout the whole Cretaceous, will have wide ranges: viz., from America through Europe to India (this is one actual case with shells in the Cretaceous period).

      LETTER 16. TO J.D. HOOKER. Down {1845}.

      I ought to have written sooner to say that I am very willing to subscribe 1 pound 1 shilling to the African man (though it be murder on a small scale), and will send you a Post-office-order payable to Kew, if you will be so good as to take charge of it. Thanks for your information about the Antarctic Zoology; I got my numbers when in Town on Thursday: would it be asking your publisher to take too much trouble to send your Botany {"Flora Antarctica," by J.D. Hooker, 1844} to the Athenaeum Club? he might send two or three numbers together. I am really ashamed to think of your having given me such a valuable work; all I can say is that I appreciate your present in two ways — as your gift, and for its great use to my species-work. I am very glad to hear that you mean to attack this subject some day. I wonder whether we shall ever be public combatants; anyhow, I congratulate myself in a most unfair advantage of you, viz., in having extracted more facts and views from you than from any one other person. I daresay your explanation of polymorphism on volcanic islands may be the right one; the reason I am curious about it is, the fact of the birds on the Galapagos being in several instances very fine-run species — that is, in comparing them, not so much one with another, as with their analogues from the continent. I have somehow felt, like you, that an alpine form of a plant is not a true variety; and yet I cannot admit that the simple fact of the cause being assignable ought to prevent its being called a variety; every variation must have some cause, so that the difference would rest on our knowledge in being able or not to assign the cause. Do you consider that a true variety should be produced by causes acting through the parent? But even taking this definition, are you sure that alpine forms are not inherited from one, two, or three generations? Now, would not this be a curious and valuable experiment (16/1. For an account of work of this character, see papers by G. Bonnier in the "Revue Generale," Volume II., 1890; "Ann. Sc. Nat." Volume XX.; "Revue Generale," Volume VII.), viz., to get seeds of some alpine plant, a little more hairy, etc., etc., than its lowland fellow, and raise seedlings at Kew: if this has not been done, could you not get it done? Have you anybody in Scotland from whom you could get the seeds?

      I have been interested by your remarks on Senecia and Gnaphalium: would it not be worth while (I should be very curious to hear the result) to make a short list of the generally considered variable or polymorphous genera, as Rosa, Salix, Rubus, etc., etc., and reflect whether such genera are generally mundane, and more especially whether they have distinct or identical (or closely allied) species in their different and distant habitats.

      Don't forget me, if you ever stumble on cases of the same species being MORE or LESS variable in different countries.

      With respect to the word "sterile" as used for male or polleniferous flowers, it has always offended my ears dreadfully; on the same principle that it would to hear a potent stallion, ram or bull called sterile, because they did not bear, as well as beget, young.

      With respect to your geological-map suggestion, I wish with all my heart I could follow it; but just reflect on the number of measurements requisite;