More Letters of Charles Darwin — Volume 2. Darwin Charles

your belief about the difficulty of dispersal of land molluscs; I was interrupted when beginning to experimentise on the just hatched young adhering to the feet of ground-roosting birds. I differ on one other point — viz. in the belief that there must have existed a Tertiary Antarctic continent, from which various forms radiated to the southern extremities of our present continents. But I could go on scribbling forever. You have written, as I believe, a grand and memorable work, which will last for years as the foundation for all future treatises on Geographical Distribution.

      P.S. — You have paid me the highest conceivable compliment, by what you say of your work in relation to my chapters on distribution in the "Origin," and I heartily thank you for it.

      LETTER 390. FROM A.R. WALLACE TO CHARLES DARWIN. The Dell, Grays, Essex, June 7th, 1876.

      Many thanks for your very kind letter. So few people will read my book at all regularly, that a criticism from one who does so will be very welcome. If, as I suppose, it is only to page 184 of Volume I. that you have read, you cannot yet quite see my conclusions on the points you refer to (land molluscs and Antarctic continent). My own conclusion fluctuated during the progress of the book, and I have, I know, occasionally used expressions (the relics of earlier ideas) which are not quite consistent with what I say further on. I am positively against any Southern continent as uniting South America with Australia or New Zealand, as you will see at Volume I., pages 398-403, and 459-66. My general conclusions as to distribution of land mollusca are at Volume II., pages 522-9. (390/1. "Geographical Distribution" II., pages 524, 525. Mr. Wallace points out that "hardly a small island on the globe but has some land-shells peculiar to it" — and he goes so far as to say that probably air-breathing mollusca have been chiefly distributed by air- or water-carriage, rather than by voluntary dispersal on the land.) When you have read these passages, and looked at the general facts which lead to them, I shall be glad to hear if you still differ from me.

      Though, of course, present results as to the origin and migrations of genera of mammals will have to be modified owing to new discoveries, I cannot help thinking that much will remain unaffected, because in all geographical and geological discoveries the great outlines are soon reached, the details alone remain to be modified. I also think much of the geological evidence is now so accordant with, and explanatory of, Geographical Distribution, that it is prima facie correct in outline. Nevertheless, such vast masses of new facts will come out in the next few years that I quite dread the labour of incorporating them in a new edition.

      I hope your health is improved; and when, quite at your leisure, you have waded through my book, I trust you will again let me have a few lines of friendly criticism and advice.

      LETTER 391. TO A.R. WALLACE. Down, June 17th, 1876.

      I have now finished the whole of Volume I., with the same interest and admiration as before; and I am convinced that my judgment was right and that it is a memorable book, the basis of all future work on the subject. I have nothing particular to say, but perhaps you would like to hear my impressions on two or three points. Nothing has struck me more than the admirable and convincing manner in which you treat Java. To allude to a very trifling point, it is capital about the unadorned head of the Argus-pheasant. (391/1. See "Descent of Man," Edition I., pages 90 and 143, for drawings of the Argus pheasant and its markings. The ocelli on the wing feathers were favourite objects of Mr. Darwin, and sometimes formed the subject of the little lectures which on rare occasions he would give to a visitor interested in Natural History. In Mr. Wallace's book the meaning of the ocelli comes in by the way, in the explanation of Plate IX., "A Malayan Forest with some of its peculiar Birds." Mr. Wallace (volume i., page 340) points out that the head of the Argus pheasant is, during the display of the wings, concealed from the view of a spectator in front, and this accounts for the absence of bright colour on the head — a most unusual point in a pheasant. The case is described as a "remarkable confirmation of Mr. Darwin's views, that gaily coloured plumes are developed in the male bird for the purpose of attractive display." For the difference of opinion between the two naturalists on the broad question of coloration see "Life and Letters," III., page 123. See Letters 440-453.) How plain a thing is, when it is once pointed out! What a wonderful case is that of Celebes: I am glad that you have slightly modified your views with respect to Africa. (391/2. "I think this must refer to the following passage in 'Geog. Dist. of Animals,' Volume I., pages 286-7. 'At this period (Miocene) Madagascar was no doubt united with Africa, and helped to form a great southern continent which must at one time have extended eastward as far as Southern India and Ceylon; and over the whole of this the lemurine type no doubt prevailed.' At the time this was written I had not paid so much attention to islands, and in my "Island Life" I have given ample reasons for my belief that the evidence of extinct animals does not require any direct connection between Southern India and Africa." — Note by Mr. Wallace.) And this leads me to say that I cannot swallow the so-called continent of Lemuria — i.e., the direct connection of Africa and Ceylon. (391/3. See "Geographical Distribution," I., page 76. The name Lemuria was proposed by Mr. Sclater for an imaginary submerged continent extending from Madagascar to Ceylon and Sumatra. Mr. Wallace points out that if we confine ourselves to facts Lemuria is reduced to Madagascar, which he makes a subdivision of the Ethiopian Region.) The facts do not seem to me many and strong enough to justify so immense a change of level. Moreover, Mauritius and the other islands appear to me oceanic in character. But do not suppose that I place my judgment on this subject on a level with yours. A wonderfully good paper was published about a year ago on India, in the "Geological Journal," I think by Blanford. (391/4. H.F. Blanford "On the Age and Correlations of the Plant-bearing Series of India and the Former Existence of an Indo-Oceanic Continent" ("Quart. Journ. Geol. Soc." XXXI., 1875, page 519). The name Gondwana-Land was subsequently suggested by Professor Suess for this Indo-Oceanic continent. Since the publication of Blanford's paper, much literature has appeared dealing with the evidence furnished by fossil plants, etc., in favour of the existence of a vast southern continent.) Ramsay agreed with me that it was one of the best published for a long time. The author shows that India has been a continent with enormous fresh-water lakes, from the Permian period to the present day. If I remember right, he believes in a former connection with S. Africa.

      I am sure that I read, some twenty to thirty years ago in a French journal, an account of teeth of Mastodon found in Timor; but the statement may have been an error. (391/5. In a letter to Falconer (Letter 155), January 5th, 1863, Darwin refers to the supposed occurrence of Mastodon as having been "smashed" by Falconer.)

      With respect to what you say about the colonising of New Zealand, I somewhere have an account of a frog frozen in the ice of a Swiss glacier, and which revived when thawed. I may add that there is an Indian toad which can resist salt-water and haunts the seaside. Nothing ever astonished me more than the case of the Galaxias; but it does not seem known whether it may not be a migratory fish like the salmon. (391/6. The only genus of the Galaxidae, a family of fresh-water fishes occurring in New Zealand, Tasmania, and Tierra del Fuego, ranging north as far as Queensland and Chile (Wallace's "Geographical Distribution," II., page 448).)

      LETTER 392. TO A.R. WALLACE. Down, June 25th, 1876.

      I have been able to read rather more quickly of late, and have finished your book. I have not much to say. Your careful account of the temperate parts of South America interested me much, and all the more from knowing something of the country. I like also much the general remarks towards the end of the volume on the land molluscs. Now for a few criticisms.

      Page 122. (392/1. The pages refer to Volume II. of Wallace's "Geographical Distribution.") — I am surprised at your saying that "during the whole Tertiary period North America was zoologically far more strongly contrasted with South America than it is now." But we know hardly anything of the latter except during the Pliocene period; and then the mastodon, horse, several great edentata, etc., etc., were common to the north and south. If you are right, I erred greatly in my "Journal," where I insisted on the former close connection between the two.

      Page 252 and elsewhere. — I agree thoroughly with the general principle that a great area with many competing forms is necessary for much and high development; but do you not extend this principle too far — I should say much too far, considering how often several species of the same genus have been developed on very small islands?

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