The Malay Archipelago, Volume 1. Alfred Russel Wallace

numerous varieties of squirrels are constantly met with. In the latter none of these occur; but the prehensile-tailed Cuscus is almost the only terrestrial mammal seen, except wild pigs, which are found in all the islands, and deer (which have probably been recently introduced) in Celebes and the Moluccas. The birds which are most abundant in the Western Islands are woodpeckers, barbets, trogons, fruit-thrushes, and leaf-thrushes; they are seen daily, and form the great ornithological features of the country. In the Eastern Islands these are absolutely unknown, honeysuckers and small lories being the most common birds, so that the naturalist feels himself in a new world, and can hardly realize that he has passed from the one region to the other in a few days, without ever being out of sight of land.

      The inference that we must draw from these facts is, undoubtedly, that the whole of the islands eastwards beyond Java and Borneo do essentially form a part of a former Australian or Pacific continent, although some of them may never have been actually joined to it. This continent must have been broken up not only before the Western Islands were separated from Asia, but probably before the extreme southeastern portion of Asia was raised above the waters of the ocean; for a great part of the land of Borneo and Java is known to be geologically of quite recent formation, while the very great difference of species, and in many cases of genera also, between the productions of the Eastern Malay Islands and Australia, as well as the great depth of the sea now separating them, all point to a comparatively long period of isolation.

      It is interesting to observe among the islands themselves how a shallow sea always intimates a recent land connexion. The Aru Islands, Mysol, and Waigiou, as well as Jobie, agree with New Guinea in their species of mammalia and birds much more closely than they do with the Moluccas, and we find that they are all united to New Guinea by a shallow sea. In fact, the 100-fathom line round New Guinea marks out accurately the range of the true Paradise birds.

      It is further to be noted—and this is a very interesting point in connection with theories of the dependence of special forms of life on external conditions—that this division of the Archipelago into two regions characterised by a striking diversity in their natural productions does not in any way correspond to the main physical or climatal divisions of the surface. The great volcanic chain runs through both parts, and appears to produce no effect in assimilating their productions. Borneo closely resembles New Guinea not only in its vast size and its freedom from volcanoes, but in its variety of geological structure, its uniformity of climate, and the general aspect of the forest vegetation that clothes its surface. The Moluccas are the counterpart of the Philippines in their volcanic structure, their extreme fertility, their luxuriant forests, and their frequent earthquakes; and Bali with the east end of Java has a climate almost as dry and a soil almost as arid as that of Timor. Yet between these corresponding groups of islands, constructed as it were after the same pattern, subjected to the same climate, and bathed by the same oceans, there exists the greatest possible contrast when we compare their animal productions. Nowhere does the ancient doctrine—that differences or similarities in the various forms of life that inhabit different countries are due to corresponding physical differences or similarities in the countries themselves—meet with so direct and palpable a contradiction. Borneo and New Guinea, as alike physically as two distinct countries can be, are zoologically wide as the poles asunder; while Australia, with its dry winds, its open plains, its stony deserts, and its temperate climate, yet produces birds and quadrupeds which are closely related to those inhabiting the hot damp luxuriant forests, which everywhere clothe the plains and mountains of New Guinea.

      In order to illustrate more clearly the means by which I suppose this great contrast has been brought about, let us consider what would occur if two strongly contrasted divisions of the earth were, by natural means, brought into proximity. No two parts of the world differ so radically in their productions as Asia and Australia, but the difference between Africa and South America is also very great, and these two regions will well serve to illustrate the question we are considering. On the one side we have baboons, lions, elephants, buffaloes, and giraffes; on the other spider-monkeys, pumas, tapirs, anteaters, and sloths; while among birds, the hornbills, turacos, orioles, and honeysuckers of Africa contrast strongly with the toucans, macaws, chatterers, and hummingbirds of America.

      Now let us endeavour to imagine (what it is very probable may occur in future ages) that a slow upheaval of the bed of the Atlantic should take place, while at the same time earthquake-shocks and volcanic action on the land should cause increased volumes of sediment to be poured down by the rivers, so that the two continents should gradually spread out by the addition of newly-formed lands, and thus reduce the Atlantic which now separates them, to an arm of the sea a few hundred miles wide. At the same time we may suppose islands to be upheaved in mid-channel; and, as the subterranean forces varied in intensity, and shifted their points of greatest action, these islands would sometimes become connected with the land on one side or other of the strait, and at other times again be separated from it. Several islands would at one time be joined together, at another would be broken up again, until at last, after many long ages of such intermittent action, we might have an irregular archipelago of islands filling up the ocean channel of the Atlantic, in whose appearance and arrangement we could discover nothing to tell us which had been connected with Africa and which with America. The animals and plants inhabiting these islands would, however, certainly reveal this portion of their former history. On those islands which had ever formed a part of the South American continent, we should be sure to find such common birds as chatterers and toucans and hummingbirds, and some of the peculiar American quadrupeds; while on those which had been separated from Africa, hornbills, orioles, and honeysuckers would as certainly be found. Some portion of the upraised land might at different times have had a temporary connection with both continents, and would then contain a certain amount of mixture in its living inhabitants. Such seems to have been the case with the islands of Celebes and the Philippines. Other islands, again, though in such close proximity as Bali and Lombock, might each exhibit an almost unmixed sample of the productions of the continents of which they had directly or indirectly once formed a part.

      In the Malay Archipelago we have, I believe, a case exactly parallel to that which I have here supposed. We have indications of a vast continent, with a peculiar fauna and flora having been gradually and irregularly broken up; the island of Celebes probably marking its furthest westward extension, beyond which was a wide ocean. At the same time Asia appears to have been extending its limits in a southeast direction, first in an unbroken mass, then separated into islands as we now see it, and almost coming into actual contact with the scattered fragments of the great southern land.

      From this outline of the subject, it will be evident how important an adjunct Natural History is to Geology; not only in interpreting the fragments of extinct animals found in the earth's crust, but in determining past changes in the surface which have left no geological record. It is certainly a wonderful and unexpected fact that an accurate knowledge of the distribution of birds and insects should enable us to map out lands and continents which disappeared beneath the ocean long before the earliest traditions of the human race. Wherever the geologist can explore the earth's surface, he can read much of its past history, and can determine approximately its latest movements above and below the sea-level; but wherever oceans and seas now extend, he can do nothing but speculate on the very limited data afforded by the depth of the waters. Here the naturalist steps in, and enables him to fill up this great gap in the past history of the earth.

      One of the chief objects of my travels was to obtain evidence of this nature; and my search after such evidence has been rewarded by great success, so that I have been able to trace out with some probability the past changes which one of the most interesting parts of the earth has undergone. It may be thought that the facts and generalizations here given would have been more appropriately placed at the end rather than at the beginning of a narrative of the travels which supplied the facts. In some cases this might be so, but I have found it impossible to give such an account as I desire of the natural history of the numerous islands and groups of islands in the Archipelago, without constant reference to these generalizations which add so much to their interest. Having given this general sketch of the subject, I shall be able to show how the same principles can be applied to the individual islands of a group, as to the whole Archipelago; and thereby make my account of the many new and curious animals which inhabit them both, more interesting and more instructive than if treated as mere isolated facts.

      Contrasts