Gulls. Professor John C. Coulson

of controversy, which include taxonomy, conservation, and the ability to identify some closely related species and subspecies in the field. Ornithology has thrived on controversy in the past and it will continue to do so in the future. In many cases, disagreement has led to the development of new methods of study and more in-depth investigations. In writing this book, debate has reared its head about where a species is placed in red or yellow categories of conservation concern based on a series of possible criteria, the existence of any one of which is enough to indicate that it is threatened. Some ornithologists feel that the system should be improved, and that a more critical scientific approach is needed to interpret data used to estimate the current risk of extinction of individual gull species.

      For the same reason, there is no comment on climate change in this book. While there have been several claims that this has affected gulls, sound evidence in favour of such effects is currently poor and is not supported by evidence that adequately allows other possible causes to be excluded. Increasing studies have been made to estimate the potential risks to gulls from offshore wind farms and the rotating arms of the turbines. At present, most of these are based on informed speculation and such factors as the flight height of individual species and their numbers in the key areas. In the future, this problem will be investigated in more detail and the level of the perceived risk to gulls and other seabirds will be based on actual information, not just models of the situation, but the research is not yet at this stage.

      In recent years, the common names of some gulls have lengthened, allegedly to avoid international confusion. Hence, the Black-legged Kittiwake, European Herring Gull, American Herring Gull and Yellow-legged Gull join the Lesser Black-backed Gull, Great Black-backed Gull, Black-headed Gull and Slender-billed Gull, which already have long names. Proposals have been made to change the Great Black-headed Gull to the shorter Pallas’s Gull, while it has been suggested that the Common Gull is changed to the Mew Gull. In this book, as mentioned earlier, Herring Gull is used for the European Herring Gull and the American Herring Gull is referred to in full. Similarly, and as mentioned above, the Black-legged Kittiwake is referred to as the Kittiwake, while its sibling species, the Red-legged Kittiwake (which is infrequently mentioned) is written out in full. In most cases this follows the vernacular English names listed in the ninth edition of ‘The Simple British List Based on a Checklist of Birds of Britain’ (2018). In general, this edition retained names already familiar to most readers.

      Statistical tests are important in evaluating differences in quantitative data, but to many the presentation of these and their outcomes are but an irritation. In general, I have commented on quantitative differences only when they have been shown to be statistically significant and so are likely to be real and meaningful, although I have not given details of the tests used.

      ACKNOWLEDGEMENTS

      Much of the information in this book derives from two sources. Some comes from professional research, but much is the result of the activities of amateur birdwatchers who spend their spare time visiting sites where they are likely to encounter unusual bird species or birds in exceptionally large numbers, and who then send the details to local recorders or contribute to national surveys and national censuses. Over the years, the number of observers has increased markedly and systems of notifying others of the presence of unusual birds have developed. Both categories of people studying birds have swelled progressively over the past 60 years, and the information and numbers of records have increased to a remarkable extent. In addition, the methods of identifying and confirming rare species have been increasingly supported by good-quality photographs. Accompanying these trends has been a dramatic increase in the numbers of gulls ringed both in Britain and Ireland, and also elsewhere in Europe. Capturing and ringing adult gulls has increased dramatically in recent years with the development of cannon nets and more frequent visits to landfill sites by teams of ringers. All of these additional efforts are appreciated, for without them, much of the information in this book would not have been available.

      I would like to acknowledge the detailed contribution of Alan Dean, both for his detailed analysis of the records of gulls in the West Midlands in England over many years and for the photographs he (and others) have so willingly contributed.

      The data used in writing this book has been collected over many years and it is possible here to identify only a small number of the hundreds of contributors. The many research students I supervised and advised have all made appreciable contributions over the years, and many have since made further contributions in ornithology and science in general while holding permanent posts both in the UK and North America. I was fortunate in having such an able set of students who contributed wholeheartedly and consistently to studies often made under difficult conditions. They all willingly volunteered to take part in teamwork as required, often at the most unsocial hours of the day and in adverse weather conditions, all while advancing their own studies.

      I owe a great deal to the BTO, not only for the ringing scheme they manage so efficiently, but also for the extensive data they have collected. When I first had contact with the BTO in the early 1950s, it had a small office at the top of a set of outside stairs in a side street in Oxford, with a single salaried member of staff, Dr Bruce Campbell. Here, I acknowledge the trust’s willingness to allow me to reproduce maps of ringing recoveries of gulls and to use the most recently available maps of the breeding season distribution of gulls in Britain and Ireland in 2007–11.

      Many people have offered photographs for inclusion in the book, and these have all been credited in the captions. In particular, I thank Nicholas Aebischer, Pep Arcos, Rob Barrett, Colin Carter, Becky Coulson, Anthony Davison, Alan Dean, Andrew Easton, Phil Jones, John Kemp, Mark Leitch, Fred van Olphen, Daniel Oro, Mike Osborne, Viola Ross-Smith, Steven Seal, Charles Sharp, Michael Southcott, Brett Spencer, Norman Deans van Swelm, Thermos (fi.wikipedia), Dan Turner and an anonymous photographer for their photographic help and willing permission to allow their excellent images to be included in this book.

      The extensive data set on British seabirds now archived and maintained by the Joint Nature Conservancy Council and readily accessible online is a valuable asset and source of information. My thanks also go to Natural England for granting a freedom of information request concerning culls and licences issued to collect gull eggs.

      Help and information have been supplied by many, in particular Nicholas Aebischer, David Baines, Robert Barrett, Peter Bell, Richard Bevan, Tim Birkhead, Bill Bourne, Joanna Burger, David Cabot, Kees Camphuysen, Colin Carter, Keith Clarkson, Ian Court, J. B. Cragg, Francis Daunt, Ian Deans, Greg Douglas, Andy Douse, Steve Dudley, Tim Dunn, George Dunnet, Andrew Easton, Julie Ellis, Mike Erwin, Sheila Frazer, Bob Furness, Michael Gochfeld, Thalassa Hamilton, Gill Hartley, Scott Hatch, Martin Heubeck, Grace Hickling, Keith Houston, Jon Jonsson, Heather Kyle, Susan Lindsay, Roddy Mavor, Jim Mills, Ian Nisbet, Daniel Oro, Ian Patterson, Ray Pierotti, Jean-Marc Pons, Julie Porter, Dick Potts, Richard Procter, Chris Redfern, Jim Reid, Sam Rider, Peter Rock, Robin Sellars, Peter Shield, Robert Swann, Mike Swindells, Martin Taylor, Mike Toms, Andrew Tongue, Mike Trees, Daniel Turner, Sarah Wanless, Matt Wood, Vero Wynne-Edwards, Bernard Zonfrillo and Jan Zorgdrager. Over the years, many others have helped in studies and investigations, and I appreciate the assistance of them all.

      I owe a major debt of gratitude to my wife, Becky, for the many ways she has supported and helped my gull studies, solved many computing problems, and given help in preparing and checking the text of this book.

       CHAPTER 1

       An Overview of Gulls

      GULLS ARE CONSPICUOUS WEB-FOOTED, long-winged, medium or large seabirds that are readily recognised by the public. Adults are mainly white with shades of grey or black on the mantle and wings. Most species have black wing-tips, some have white ‘mirrors’ within the black areas, but a few species – mainly those restricted to an Arctic breeding distribution – have entirely white wing-tips. In the breeding season, adults of different species either have entirely pure white or very dark (black or brown) heads, and all revert to white heads in the autumn and winter, often with small grey marks behind the eye or grey streaking on the neck.

      Gulls are widely